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They are likely to pay higher fees compared to other services/wallets that have direct channels to the destinations they want to reach. To pay a CashApp user it's also very likely that you have to pay more compared to other services.
The c= node adds an extra hop to every route that goes from/to the CashApp nodes. Fees c= charges are 1000 to 3000 ppm, so 0.1 to 0.3 percent is added to each payment.
Since most of the volume CashApp produces is sending, the c= node gets outbound heavy so they have to close the channels that the CashApp nodes open. They use the funds from those closed channels to open new channels to the destinations the CashApp users want to reach. This is how they are able to earn so much from "routing".
Originally CashApp did only peer with select nodes because of legal and compliance reasons. I guess at some point they figured they could earn a bunch if they put a node running LDK between them and the rest of the network. It's frankly a bit strange how they act like they earn from routing while they are just an exclusive gatekeeper for CashApp lightning.